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45. Burraco, P., Salla, R. and Orizaola, G. 2023. Exposure to ionizing radiation and liver histopathology in the tree frogs of Chornobyl (Ukraine). Chemosphere 315, 137753. LINK


44. Burraco, P. and Orizaola, G. 2022. Ionizing radiation and melanism in Chornobyl tree frogs. Evolutionary Applications 15, 1469-1479. LINK

43. Orizaola, G., Wood, M.D., Oughton, D.H., Gashchak, S., Vishnevskiy, D. and Beresford, N.A. 2022. Ukraine: restore Chernobyl’s radioecology collaboration. Nature 605, 225. LINK

PRE-PRINT. Videvall, E., Burraco, P. and Orizaola, G. 2022. Impact of ionizing radiation on the environmental microbiomes of Chernobyl wetlands. bioXriv, 476627. LINK

42. Car, C., Gilles, A., Armant, O., Burraco, P., Beaugelin-Seiller, K., Gashchak, S., Camilleri, V., Cavalie, I., Laloi, P., Adam-Guillermin, C., Orizaola, G. and Bonzom, J.-M. 2022. Unusual evolution of tree frog populations in the Chernobyl exclusion zone. Evolutionary Applications 15, 203-219. LINK

BOOK CHAPTER. Orizaola, G. 2022. Amphibians in field radioecology: a review and perspective. In: Biomarkers of radiation in the environment – Robust tools for risk assessment (eds. M. D. Wood, C. E. Mothersill, G. Tsakanova, T. Cresswell, and G. E. Woloschak). NATO Science for Peace and Security Series A: Chemistry and Biology. Springer, pp. 185-203.


41. Burraco, P., Car, C., Bonzom, J.-M. and Orizaola, G. 2021. Assessment of exposure to ionizing radiation in Chernobyl tree frogs (Hyla orientalis). Scientific Reports 11, 20509

40. Burraco, P., Bonzom, J.-M., Car, C., Beaugelin-Seiller, K., Gashchak, S. and Orizaola, G. 2021. Lack of impact of radiation on blood physiology biomarkers of Chernobyl tree frogs. Frontiers in Zoology 18, 33. LINK

39. Burraco, P., Laurila, A. and Orizaola, G. 2021. Limits to compensatory responses to altered phenology in amphibian larvae. Oikos 130, 231-239. LINK

BOOK CHAPTER. Orizaola, G. 2021. Chernobyl has become a refuge for wildlife thirty-three years after the nuclear accident. In: No, you’re not entitled to your opinion, and 49 other essays that got the world talking (A. Hansen, ed.). Thames & Hudson / The Conversation, pp. 38-41.


38. Burraco, P., Orizaola, G., Monaghan, P. and Metcalfe, N. 2020. Climate change and ageing in ectotherms. Global Change Biology 26, 5371-5281. LINK.

37. Johansson, F., Orizaola, G. and Nilsson-Örtman, V. 2020. Temperate insects with narrow seasonal activity periods can be as vulnerable to climate change as tropical insect species. Scientific Reports 10, 8822. LINK

36. Burraco, P., Valdés, A. E. and Orizaola, G. 2020. Metabolic costs of altered growth trajectories across life transitions in amphibians. Journal of Animal Ecology 89, 855-866. LINK

35. Beresford, N., Horemans, N., Copplestone, D., Raines, K.E., Orizaola, G., Wood, M.D., Laanen, P., Whitehead, H.C., Burrows, J.E., Tinsley, M.C., Smith, J.T., Bonzom, J.-M., Gagnaire, B., Adam-Guillermin, C., Gashchak, S., Jha, A.N., de Menezes, A., Willey, N. and Spurgeon, D. 2020. Towards solving a scientific controversy – The effects of ionising radiation on the environment. Journal of Environmental Radioactivity 211, 106033. LINK

34. Orizaola, G. 2020. From nuclear desert to evolutionary lab: the response of living organisms to Chernobyl’s ionising radiation. Mètode Science Studies Journal 10, 193-199. LINK


33. Lindgren, B., Orizaola, G. and Laurila, A. 2018. Interacting effects of predation risk and resource level on escape speed of amphibian larvae along a latitudinal gradient. Journal of Evolutionary Biology 31, 1216-1226. LINK


32. Murillo-Rincón, A.P., Laurila, A. and Orizaola, G. 2017. Compensating for delayed hatching reduces offspring immune response and increases life-history costs. Oikos 126, 565-571. LINK

31. Murillo-Rincón, A.P., Kolter, N. A., Laurila, A. and Orizaola, G. 2017.  Intra-specific priority effects modify compensatory responses to changes in hatching phenology in an amphibian. Journal of Animal Ecology 86, 128-135. LINK 


30. Carreira, B., Segurado, P., Orizaola, G., Gonçalves, N.,  Pinto, V., Laurila, A. and Rebelo, R. 2016. Warm vegetarians? Heat waves and diet shifts in tadpoles. Ecology 97, 2464-2474. LINK 

Photo Gallery at ESA Bulletin 98, 53-56 PDF

29. Orizaola, G., Richter-Boix, A. and Laurila, A. 2016. Transgenerational effects and impact of compensatory responses to changes in breeding phenology on antipredator defenses. Ecology 97, 2470-2478. LINK

Photo Galley at ESA Bulletin 98, 47-52 PDF

28. Orizaola, G. and Laurila, A. 2016. Developmental plasticity Issue Informationincreases at the northern range margin in a warm-dependent amphibian. Evolutionary Applications 9, 471-478. LINK (Open Access).


27. Dijk, B., Laurila, A., Orizaola, G. and Johansson, F. 2016. Is one defence enough? Disentangling the relative importance of morphological and behavioural predator induced defences. Behavioral Ecology and Sociobiology 70, 237-246. LINK


26. Orizaola, G. and Valdés, A.E. 2015. Free the tweet at scientific conferences. Science 350, 170-171. LINK


25. Richter-Boix, A., Orizaola, G. and Laurila, A. 2014. Transgenerational phenotypic plasticity links parental phenology with offspring life-history. Ecology, 2715-2722. LINK

24. Nunes, A.L., Orizaola, G., Laurila, A. and Rebelo, R. 2014. Rapid evolution of constitutive and inducible defenses against an invasive predator. Ecology 95, 1520-1530. PDF (pre-print)

Photo Galley at ESA Bulletin 95, 451- 456 PDF

23.  Nunes, A.L., Orizaola, G., Laurila, A. and Rebelo, R. 2014. Morphological and life-history responses of anurans to an invasive predator: an integrative approach. Ecology and Evolution 4, 1491-1503 (Open Access LINK).

22. Orizaola, G., Dahl, E. and Laurila, A. 2014. Compensatory growth strategies are affected by the strength of environmental time-constraints in anuran larvae. Oecologia 174, 131-137. PDF


21. Orizaola, G., Dahl, E., Nicieza, A.G. and Laurila, A. 2013. Life history and anti-predator strategies are affected by breeding phenology in an amphibian. Oecologia 171, 873-881. PDF


20. Dahl, E., Orizaola, G., Nicieza, A.G. and Laurila, A. 2012. Time constraints and flexibility of growth strategies: geographic variation in catch-up growth responses in amphibian larvae. Journal of Animal Ecology 81, 1233-1243. PDF

19. Dahl, E., Orizaola, G., Windberg, S. and Laurila, A. 2012. Geographic variation in corticosterone response to chronic predator stress in tadpoles. Journal of Evolutionary Biology 25, 1066-1076. PDF

18. Orizaola, G., Dahl, E. and Laurila, A. 2012. Reversibility of predation-induced plasticity and its effect at a life history switch point. Oikos 121, 44-52. PDF


17. Arce, F., Orizaola, G. and Navedo, J.G. 2011. Storm petrel´s breeding skipping in response to oil-spill pollution: Raising concerns over Zabala et al. (2011) methodological approach. Marine Pollution Bulletin 62, 2576-2577. PDF


16. Tejedo, M., Marangoni, F., Pertoldi, C., Richter-Boix, A., Laurila, A., Orizaola, G., Nicieza, A.G., Álvarez, D. and Gómez-Mestre, I. 2010. Contrasting effects of environmental factors during larval stage on morphological plasticity in post-metamorphic frogs. Climate Research 41, 31-39. PDF

15. Navedo, J.G., Orizaola, G., Masero, J.A., Overdijk, O. and Sánchez-Guzmán, J.M. 2010. Long-distance travellers stopover for longer: a case study with spoonbills staying in North Iberia. Journal of Ornithology, 151, 915-921. PDF

14. Orizaola, G., Quintela, M. and Laurila, A. 2010. Climatic adaptation in an isolated and genetically impoverished amphibian population. Ecography 33, 730-737. PDF

13. Orizaola, G., Dahl, E. and Laurila, A. 2010. Compensating for delayed hatching across consecutive life-history stages in an amphibian. Oikos 119, 980-987. PDF

12. Navedo, J.G., Masero, J.A., Overdijk, O., Orizaola, G. and Sánchez-Guzmán, J.M. 2010. Assessing the role of multiple environmental factors on Eurasian Spoonbill departure decisions from stopover sites. Ardea, 98, 3-12. PDF


11. Orizaola, G. and Laurila, A. 2009. Microgeographic variation in temperature-induced plasticity in an isolated amphibian metapopulation. Evolutionary Ecology 26, 979-991. PDF

10. Orizaola, G. and Laurila, A. 2009. Intraspecific variation of temperature-induced effects on metamorphosis in the pool frog (Rana lessonae). Canadian Journal of Zoology 87, 581-588. PDF

9. Orizaola, G. and Laurila, A. 2009. Microgeographic variation in the effects of larval temperature environment on juvenile morphology and locomotion in the pool frog. Journal of Zoology 277, 267-274. PDF


8. Orizaola, G. and Laurila, A. 2008. Growing with kin does not bring benefits to tadpoles in a genetically impoverished amphibian population. Canadian Journal of Zoology 86, 45-50. PDF


7. Orizaola, G. and Braña, F. 2006. Effect of salmonid introduction and other environmental characteristics on amphibian distribution and abundance in mountain lakes of Northern Spain. Animal Conservation 9, 171-178. PDF


6. Orizaola, G. and Braña, F. 2005. Plasticity in newt metamorphosis: the effect of predation at embryonic and larval stages. Freshwater Biology 50, 438-446. PDF *Front cover article*


5. Orizaola, G. and Braña, F. 2004. Hatching responses of four newt species to predatory fish chemical cues. Annales Zoologici Fennici 41, 635-645. PDF


4. Orizaola, G. and Braña, F. 2003. Response of predator-naive newt larvae to food and predator presence. Canadian Journal of Zoology 81, 1845-1850. PDF

3. Orizaola, G. and Braña, F. 2003. Do predator chemical cues affect newt oviposition site selection? Herpetological Journal 13, 189-193. PDF

2. Orizaola, G. and Braña, F. 2003. Oviposition behaviour and vulnerability of eggs to predation in four newt species (genus Triturus). Herpetological Journal 13, 121-124. PDF


1. Braña, F., Frechilla, L. and Orizaola, G. 1996. Effect of introduced fish on amphibian assemblages in mountain lakes of northern Spain. Herpetological Journal 6, 145-148. PDF